The upward tilt of honeycomb cells increases the carrying capacity of the comb and is not to prevent the outflow of honey
Crystallography of honeycomb formation under geometric frustration
A simplified thermomechanical approach to visualize hexagonal honeycomb construction
Hexagons: The structural possibilities of a uniform shape
How do bees produce the hexagonal-shaped cells of a honeycomb? - Quora
Why do bees make hexagons in their hives and not squares, rectangles, or triangles? - Quora
Honeybee combs: how the circular cells transform into rounded hexagons
Honeycomb - Wikipedia
How do bees produce the hexagonal-shaped cells of a honeycomb? - Quora
The bee Tetragonula builds its comb like a crystal
Why do bees make hexagons in their hives and not squares, rectangles, or triangles? - Quora
Imperfect comb construction reveals the architectural abilities of honeybees